as the root grow, so the chambers function changes and new chambers built, ones with a specific function and others from roots intersections.
the general position of the ants stay is basically divided to 3 in vertical flow.
the lower or the deeper part of the nest is the position of the birth givers and the eggs, according to researches for two reasons.keeping them safe and for a lower temperature.
other behavior of those chamber is that there is always more than one root getting there keeping the air flow. other function in the lower parts is the garbage chambers there most of the things that was gathered and not needed are kept there most of those chambers have only one way and the root there is very long .
the middle part is the position of the food storage and the staying of the broad workers , most of the expansion happening in this area merging between chambers and building new roots, in fact the connection between chambers some of the time happening from both dig direction and the position of the opposite ant discovered by the pheromone spread  and than they can control complex root.

the upper part which more dense than others, is the area of the collectors and the search groups, this part most of the time half empty and doesn't really expand after the first years of the nest. the chambers are bigger and characterized with lots of root intersection that helpfull for larger movement to the lower part of the nest and carrying object that eventually kept in the storage or the garbage.

with this information we can tell that the architecture of the nest Determined by the main roots and the chambers are divided to 2 , esential which th main root conect between , and the intoative which built between those stratagys.